Tuesday, May 24, 2011

Fish

A fish is any gill-bearing aquatic vertebrate  animal that lacks limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish, as well as various extinct related groups. Because the term is defined negatively, and excludes the tetrapods  which descend from within the same ancestry, it is paraphyletic. The traditional term pisces  is considered a typological, but not a phylogenetic classification.
Most fish are "cold-blooded", or ectothermic, allowing their body temperatures to vary as ambient temperatures change. Fish are abundant in most bodies of water. They can be found in nearly all aquatic environments, from high mountain streams  to the abyssal and even hadal depths of the deepest oceans . At 32,000 species, fish exhibit greater species diversity than any other class of vertebrates.
Fish, especially as food, are an important resource worldwide. Commercial and subsistence fishers hunt fish in wild fisheries  or farm them in ponds or in cages in the ocean . They are also caught by recreational fishers, kept as pets, raised by fishkeepers, and exhibited in public aquaria. Fish have had a role in culture through the ages, serving as deities, religious symbols, and as the subjects of art, books and movies.

     
The term "fish" most precisely describes any non-tetrapod craniate that has gills throughout life and whose limbs, if any, are in the shape of fins. Unlike groupings such as birds or mammals, fish are not a single clade but a paraphyletic collection of taxa, including hagfishes, lampreys, sharks and rays, ray-finned fish, coelacanths, and lungfish. Indeed, lungfish and coelacanths are closer relatives of tetrapods  than of other fish such as ray-finned fish or sharks, so the last common ancestor of all fish is also an ancestor to tetrapods. As paraphyletic groups are no longer recognised in modern systematic biology, the use of the term "fish" as a biological group must be avoided.
Many types of aquatic animals commonly referred to as "fish" are not fish in the sense given above; examples include shellfish, cuttlefish, starfish, crayfish and jellyfish. In earlier times, even biologists did not make a distinction – sixteenth century natural historians classified also seals, whales, amphibians, crocodiles, even hippopotamuses, as well as a host of aquatic invertebrates, as fish. However, according the definition above, all mammals, including Cetaceans like Whales and Dolphins, are not fish. In some contexts, especially in aquaculture, the true fish are referred to as fin fish  to distinguish them from these other animals.
A typical fish is ectothermic, has a streamlined body for rapid swimming, extracts oxygen from water using gills or uses an accessory breathing organ to breathe atmospheric oxygen, has two sets of paired fins, usually one or two  dorsal fins, an anal fin, and a tail fin, has jaws, has skin that is usually covered with scales, and lays eggs.
Each criterion has exceptions. Tuna, swordfish, and some species of sharks show some warm-blooded adaptations—they can heat their bodies significantly above ambient water temperature. Streamlining and swimming performance varies from fish such as tuna, salmon, and jacks that can cover 10–20 body-lengths per second to species such as eels and rays that swim no more than 0.5 body-lengths per second. Many groups of freshwater fish extract oxygen from the air as well as from the water using a variety of different structures. Lungfish have paired lungs similar to those of tetrapods, gouramis have a structure called the labyrinth organ that performs a similar function, while many catfish, such as Corydoras extract oxygen via the intestine or stomach. Body shape and the arrangement of the fins is highly variable, covering such seemingly un-fishlike forms as seahorses, pufferfish, anglerfish, and gulpers. Similarly, the surface of the skin may be naked, or covered with scales of a variety of different types usually defined as placoid , cosmoid, ganoid, cycloid, and ctenoid . There are even fish that live mostly on land. Mudskippers feed and interact with one another on mudflats and go underwater to hide in their burrows. The catfish Phreatobius cisternarum lives in underground, phreatic habitats, and a relative lives in waterlogged leaf litter.
Fish range in size from the huge 16-metre  whale shark to the tiny 8-millimetre  stout infantfish.

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